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Hmm, if I'm not mistaken an underscore is part of the 7 bit ASCII range. Even though a question mark is as well, it has the purpose to denote query strings whereas not so much the underscore. From a SEO perspective, it may be beneficial to use all hyphens as well. Matt Cutt's has also mentioned many times to use hyphens instead of underscores. (Yea, another Matt Cutt's said ) Although I suppose that would be in the sense that you should avoid underscores for all of the word separations of your content. I assume in the rare circumstances people use underscores, this would have less of an impact on SEO. I may be just a bit picky here.